1982, Bachelor in Biology, University of Lisboa
1983, MSc in Biological Oceanography (Diplome d’Études Approfondies), University Pierre et Marie Curie, Paris VI;
1988, PhD in Biological Oceanography, University Pierre et Marie Curie, Paris VI.
Spatial patterns and succession in benthic estuarine and coastal shelf systems.
Community level indicators of change of natural and anthropogenic stressors.
Integration of environmental, biological and eco-toxicological descriptors in the assessment of estuarine and coastal areas.
Characterization and classification of habitats and living resources in coastal and estuarine areas.